{"id":653367,"url":"https://alion.io/job/afterquery-senior-software-engineer-securityinfrastructure","title":"Senior Software Engineer - Security/Infrastructure","company":{"id":668886,"name":"AfterQuery","domain":"afterquery.com","url":"https://alion.io/company/afterquery","size_band":"201-500","is_staffing_agency":false,"employer_type":"direct","is_intermediary":false,"listed_via":null,"ats_vendor":"Ashby","truth_index":{"grade":"A","score":88,"open_postings":12,"ghost_share":0,"stale_share":0.5,"repost_share":0,"time_to_fill_p50_days":null,"computed_at":"2026-10-30T05:48:23Z"}},"role":"Backend","role_family":"Backend","seniority":"senior","employment_type":"full_time","work_mode":"on_site","remote_scope":null,"remote_scope_basis":null,"remote_working_hours":null,"hiring_geo_confidence":"structured","locations":["San Francisco, United States"],"countries":["US"],"hiring_countries":[],"hiring_countries_total":0,"salary":{"min":200000,"max":350000,"currency":"USD","period":"year","gross":null,"usd_annual":350000},"salary_estimate":null,"experience_years_min":2,"visa_sponsorship":false,"relocation_package":false,"has_equity":false,"technologies":[{"name":"Anthropic","optional":false},{"name":"Human-in-the-Loop","optional":false},{"name":"IAM","optional":false},{"name":"OpenAI","optional":false},{"name":"Python","optional":false},{"name":"TypeScript","optional":false},{"name":"AWS","optional":true},{"name":"GCP","optional":true}],"status":"live","first_seen_at":"2026-05-26T06:14:48Z","employer_posted_date":"2026-05-26","last_verified_at":"2026-10-31T02:10:59Z","board_verified":true,"closed_at":null,"days_open":157,"trust":{"level":"ok","repost_count":0,"flags":[],"days_open":155},"description":"About AfterQuery\nAfterQuery is an applied research lab curating data solutions for foundation model development. We serve every frontier AI lab with the mission of delivering the best data to power the best models. In doing so, we can make expertise that once took a lifetime to build available to anyone who needs it.\nOur customers are the ones building the foundation models themselves and our work sits directly in the loop of how those systems improve. This is a rare opportunity to join a company at a defining moment in AI. Forbes reported that we could be YC's fastest unicorn, reportedly raising at a $3.2 billion valuation. We're based in San Francisco and backed by leading investors including Altos Ventures, BoxGroup, and Y Combinator and angels from Google DeepMind, OpenAI, Anthropic, Meta Superintelligence Labs, and Microsoft AI.\nWhy Apply\nMassive Opportunity: Forbes reported that we could be YC's fastest unicorn, reportedly raising at a $3.2 billion valuation, and we're not slowing down.\nFounding Impact: You will own and architect core infrastructure systems that power our platform from the ground up.\nEquity & Growth: Competitive salary and meaningful equity. As we scale, you’ll have the opportunity to shape the engineering organization and lead major technical initiatives.\nStrong Team: Our founding team has experience from Citadel Securities, Meta, Google, Silver Lake, and Morgan Stanley - work alongside world-class engineers and researchers.\nOverview\nAs the Senior SWE- Security Infrastructure, you’ll be responsible for protecting a system that handles highly sensitive workflows across AI training, expert networks, and enterprise integrations with leading model labs.\nThis role is deeply technical. You’ll design and implement the infrastructure that keeps our systems resilient, rather than managing off-the-shelf security tools. We move fast and rely heavily on AI internally. You should be comfortable using LLMs to accelerate debugging, threat analysis, and system design and just as comfortable replacing repetitive work with code. Security here isn’t isolated. When you’re not working on security problems, you’ll be building core systems especially human-in-the-loop platforms and data infrastructure that power how frontier AI models are developed. We work in-person in San Francisco.\nResponsibilities\nBuild systems that identify and respond to real security events across our stack\n\nReplace manual security workflows with programmatic, scalable solutions\n\nStrengthen our cloud and production environments (GCP/AWS, containers, networking)\n\nDesign and enforce access control systems across internal teams, experts, and customers\n\nIntegrate security checks directly into development workflows without slowing shipping speed\n\nOwn incident handling systems and processes from detection through resolution\n\nSecure surfaces unique to AI systems, including data flows and model interaction layers\n\nDevelop human-in-the-loop systems used in AI training and evaluation\n\nContribute to core infrastructure and internal tooling used across the company\n\nShip features across the stack that directly impact customers and model performance\n\nRequired Qualifications\nYou’ve built internal tools or systems to solve security problems-not just configured existing products\n\nStrong programming ability in Python, Go, or TypeScript\n\nExperience working with cloud environments (IAM, networking, containers)\n\nAbility to identify vulnerabilities directly in code and system design\n\nExperience creating or maintaining signal-based monitoring and alerting systems\n\nComfortable owning production issues when they arise\n\nAble to move fluidly between security work and general engineering\n\n2-7 years building software or security systems\n\nPreferred Qualifications\nBackground in fast-scaling startups or high-output engineering teams\n\nExposure to AI systems, data pipelines, or model evaluation workflows\n\nFamiliarity with offensive security techniques\n\nExperience building tools that other engineers depend on\n\nEvidence of building and shipping systems from zero to production\n\nCompany Benefits (For Eligible Employees):\nHealth Insurance: Medical, Vision, Dental\n\n401(k) with Employer Match\n\nDaily Meals: Daily UberEats Stipend\n\nMonthly Wellness Stipend\n\nWe are an equal opportunity employer committed to providing a workplace free from discrimination and harassment. Employment decisions are made without regard to legally protected characteristics under applicable federal, state, or local law.\nWe comply with applicable pay transparency requirements and provide compensation ranges based on the position, qualifications, experience, and other relevant factors. Reasonable accommodations are available to qualified individuals with disabilities and for sincerely held religious beliefs, as required by law. This job description is intended to describe the general nature and level of work performed and is not an exhaustive list of all duties, responsibilities, qualifications, or working conditions associated with the position. 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